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Hamiltonian spite
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Hamiltonian spite : ウィキペディア英語版
Hamiltonian spite

Within the field of social evolution, Hamiltonian spite is a term for behaviours occurring among conspecifics that have a cost for the actor and a negative impact upon the recipient.
==Theories on altruism and spitefulness==

W. D. Hamilton published an influential paper on altruism in 1964 to explain why genetic kin tend to help each other.〔Hamilton WD 1964. The genetical evolution of social behaviour. Part I and II. Journal of Theoretical Biology 7: 1–16 and 17–52.〕 He argued that genetically related individuals are likely to carry the copies of the same alleles; thus, helping kin may ensure that copies of the actors' alleles pass into next generations.
While this became a widely accepted idea, it was less noted that Hamilton published a later paper that modified this view.〔Hamilton WD 1970. Selfish and spiteful behaviour in an evolutionary model. Nature 228: 1218–20.〕 This paper argues that by measuring the genetic relatedness between any two (randomly chosen) individuals of a population several times, we can identify an average level of relatedness. Theoretical models predict that (1) it is adaptive for an individual to be altruistic to any other individuals that are more closely related to it than this average level, and also that (2) it is adaptive for an individual to be spiteful against any other individuals that are less closely related to it than this average level. The indirect adaptive benefits of such acts can surpass certain costs of the act (either helpful or harmful) itself. Hamilton mentioned birds and fishes exhibiting infanticide (more specifically: ovicide) as examples for such behaviours.
Briefly, an individual can increase the chance of its genetic alleles to be passed to the next generations either by helping those that are more closely related, or by harming those that are less closely related than relationship by chance.〔Vickery WL, Brown JS, FitzGerald GJ 2003. Spite: altruism’s evil twin. Oikos, 102, 413–16.〕〔Foster KR, Wenseleers T, Ratnieks FLW 2001. Spite: Hamilton’s unproven theory. Annales Zoologici Fennici 38: 229–38.〕

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